1999年和2002年分别获北京大学学士和硕士学位，2007年获日本东京大学博士学位，2007年至2010年在日本国立环境研究所进行博士后工作，2010年至2011年在北京大学城市与环境学院任副研究员。2011年加入中国科学院植物研究所植被结构与功能研究组。

主要研究工作：

主要从事植物生理生态学、全球变化生态学和生态草牧业方面的研究，近年来重点围绕我国草原灌丛化及其对生态系统结构与功能的影响、氮沉降对北方林碳循环的影响、以及生态健康条件下的草畜平衡及其调控机制等开展研究。

主持和参加的科研项目：

1. 中国科学院战略性先导科技专项(XDA26000000)，课题“草畜动态平衡智能预测与区域模式研究”，2020-2025, 负责

2. 云南省科技厅重点研发项目(202303AC100009)，专题“云南草地碳汇潜力评估”，2023-2025，负责

3. 云南省基础研究专项重大项目，课题“云南植被资源动态监测与综合数据管理平台”，2021-2024，共同负责

4. 国家自然科学基金委基础科学中心项目(31988102)，“生态系统对全球变化的响应”，2020-2024，参加

5. 国家自然科学基金面上项目(31971747)，“灌木和草本水分利用策略的多样性及其对草原灌丛化的影响”，2020-2023，负责

6. 中国科学院前沿科学重点研究项目(QYZDY-SSW-SMC011)，“构建联接森林群落结构与物质生产的新方法和新理论”，2016-2020，参加

7. 中国科学院科技服务网络计划(KFJ-STS-ZDTP-056)，“不同区域草牧业发展的关键技术研发”，2019-2020，参加

8. 科技部基础性工作专项(2015FY210200)，专题“植被综合数据库与共享平台的构建”，2015-2020，负责

9. 国家自然科学基金面上项目(31470525)，“内蒙古草原灌丛化对土壤有机碳的影响及其驱动机制”，2015-2018，负责

10. 国家自然科学基金重点项目(31330012)，“我国温带草原的灌丛化及其对生态系统结构和功能的影响”，2014-2018，参加

11. 科技部国家重大科学研究计划(2014CB954001)，“土壤系统碳动态、机制及其对全球变化的响应”，2014-2018，参加

代表性论文（* 标记为通讯作者，^§ 为共同第一作者）：

2024

Kang J, Shen HH*, Zhang SY, Xu LC, Tang ZY, Tang YH, Fang JY. 2024. Contrasting growth responses to drought in three species widely distributed in northern China. Science of the Total Environment, 908: 168331.

Xing AJ, Shen HH*, Xu LC, Zhao MY, Yan ZB, Fang JY. 2024. Linking nutrient resorption stoichiometry with plant growth under long-term nitrogen addition. Forest Ecosystems, 11: 100221.

2023

Zhao MY, Shen HH*, Zhu YK, Xing AJ, Kang J, Liu LL, Fang JY*. 2023. Asymmetric responses of abundance and diversity of N-cycling genes to altered precipitation in arid grasslands. Functional Ecology, 37: 2953-2966.

Wang ZX, Xing AJ, Shen HH*. 2023. Effects of nitrogen addition on the combined global warming potential of three major soil greenhouse gases: A global meta-analysis. Environmental Pollution, 334: 121848.

Xing AJ, Shen HH*, Zhao MY, Fang JY. 2023. Long-term nitrogen addition reduced soil respiration contributed by decline in the understory plant cover but not species richness. Plant and Soil, 492: 125-134.

2022

Xing AJ^§, Du EZ^§, Shen HH*, Xu LC, De Vries W, Zhao MY, Liu XY, Fang JY*. 2022. Nonlinear responses of ecosystem carbon fluxes to nitrogen deposition in an old-growth boreal forest. Ecology Letters, 25: 77-88.

Xing AJ, Du EZ, Shen HH*, Xu LC, Zhao MY, Liu XY, Fang JY*. 2022. High-level nitrogen additions accelerate soil respiration reduction over time in a boreal forest. Ecology Letters, 25: 1869-1878.

Zhu YK, Shen HH*, Akinyemi DS, Zhang PJ, Feng YP, Zhao MY, Kang J, Zhao X, Hu HF, Fang JY*. 2022. Increased precipitation attenuates shrub encroachment by facilitating herbaceous growth in a Mongolian grassland. Functional Ecology, 36: 2356-2366.

Xing AJ, Xu LC, Zhao MY, Shen HH*, Ma SH, Fang JY. 2022. Shifts in understory plant composition induced by nitrogen addition predict soil fungal beta diversity in a boreal forest. Biology and Fertility of Soils, 58: 667-677.

2021

Xu LC, Xing AJ, Du EZ, Shen HH*, Yan ZB, Jiang L, Tian D, Hu HF, Fang JY. 2021. Effects of nitrogen addition on leaf nutrient stoichiometry in an old-growth boreal forest. Ecosphere, 12: e03335.

Zhu YK, Shen HH*, Feng YP, Li H, Akinyemi DS, Hu HF, Fang JY. 2021. Effects of shrub encroachment on soil aggregates and organic carbon vary in different grasslands in Inner Mongolia, China. Ecosphere, 12: e03363.

2020

Liu SS, ZhouLH, Li H, Zhao X, Yang YH, Zhu YK, Hu HF, Chen LY, Zhang PJ, Shen HH*, Fang JY*. 2020. Shrub encroachment decreases soil inorganic carbon stocks in Mongolian grasslands. Journal of Ecology, 108: 678-686.

Zhou LH^§, Liu SS^§, Shen HH*, Zhao MY, Xu LC, Xing AJ, Fang JY*. 2020. Soil extracellular enzyme activity and stoichiometry in China’s forests. Functional Ecology, 34: 1461-1471.

Akinyemi DS^§, Zhu YK^§, Zhao MY, Zhang PJ, Shen HH*, Fang JY. 2020. Response of soil extracellular enzyme activity to experimental precipitation in a shrub-encroached grassland in Inner Mongolia. Global Ecology and Conservation, 23: e01175.

2019

Li H, Shen HH*, Zhou LH, Zhu YK, Chen LY, Hu HF, Zhang PJ, Fang JY*. 2019. Shrub encroachment increases soil carbon and nitrogen stocks in temperate grasslands in china.Land Degradation and Development, 30: 756-767.

Liu SS, Shen HH*, Chen SC, Zhao X, Biswas A, Jia XL, Shi Z*, Fang JY. 2019. Estimating forest soil organic carbon content using vis-nir spectroscopy: Implications for large-scale soil carbon spectroscopic assessment. Geoderma, 348: 37-44.

Liu SS, Shen HH, Zhao X, Zhou LH, Li H, Xu LC, Xing AJ, Fang JY*. 2019. Estimation of plot-level soil carbon stocks in china's forests using intensive soil sampling. Geoderma, 348:107-114.

Xing AJ^§, Xu LC^§, Shen HH*, Du EZ, Liu XY, Fang JY. 2019. Long term effect of nitrogen addition on understory community in a Chinese boreal forest. Science of the Total Environment, 646: 989-995.

Zhou LH, Shen HH, Chen LY, Li H, Zhang PJ, Zhao X, Liu TY, Liu SS, Xing AJ, Hu HF, Fang JY*. 2019. Ecological consequences of shrub encroachment in the grasslands of northern China. Landscape Ecology, 34: 119-130.

Zhou LH, Shen HH*, Chen LY, Li H, Zhang PJ, Zhao X, Liu TY, Liu SS, Xing AJ, Hu HF, Fang JY. 2019. Species richness and composition of shrub-encroached grasslands in relation to environmental factors in northern China. Journal of Plant Ecology, 12: 56-66.

Zhou LH, Shen HH*, Xu YP, Li H, Chen LY, Wang YH, Zhang PJ, Zhao X, Liu TY, Hu HF, Fang JY*. 2019. Soil organic carbon components in Inner Mongolian shrub-encroached grasslands. Plant and Soil, 442: 199-213.

2018

Zhou LH, Li H, Shen HH, Xu YP, Wang YH, Xing AJ, Zhu YK, Zhou SZ, Fang JY*. 2018. Effects of shrub encroachment on vertical changes in soil organic carbon in Mongolian grasslands: using a multi-biomarker approach. Plant and Soil, 431: 217-230.

Liu SS, Yang YH, Shen HH, Hu HF, Zhao X, Li H, Liu TY, Fang JY*. 2018. No significant changes in topsoil carbon in the grasslands of northern China between the 1980s and 2000s. Science of the Total Environment, 624: 1478-1487.

2017

Zhou LH, Li H, Shen HH*, Xu YP, Wang YH, Xing AJ, Fang JY. 2017. Shrub-encroachment induced alteration in input chemistry and soil microbial community affect topsoil organic carbon in an Inner Mongolian grassland. Biogeochemistry, 136: 311-324.

Li H, Zhang JH, Hu HF, Chen LY, Zhu YK, Shen HH*, Fang JY*. 2017. Shift in soil microbial communities with shrub encroachment in Inner Mongolia grasslands, China. European Journal of Soil Biology, 79: 40-47.

2016

Li H, Shen HH, Chen LY, Liu TY, Hu HF, Zhao X, Zhou LH, Zhang PJ, Fang JY*. 2016. Effects of shrub encroachment on soil organic carbon in global grasslands.Scientific Reports, 6: 28974.

沈海花, 朱言坤, 赵霞, 耿晓庆, 高树琴, 方精云. 2016. 中国草地资源的现状分析. 科学通报, 61: 139-154.

2015及以前

Chen LY, Li H, Zhang PJ, Zhao X, Zhou LH, Liu TY, Hu HF, Bai YF, Shen HH*, Fang JY*. 2015. Climate and native grassland vegetation as drivers of the community structures of shrub-encroached grasslands in inner Mongolia, China. Landscape Ecology, 30:1627-1641.

陈蕾伊, 沈海花, 方精云. 2014. 灌丛化草原: 一种新的植被景观. 自然杂志, 3: 391-396.

Shen HH, Wang SP, Tang YH. 2013. Grazing alters warming effects on leaf photosynthesis and respiration in Gentiana straminea, an alpine forb species.Journal of Plant Ecology, 6: 418-427.